[1] GRANSTRÖM T B,TAKATA G,TOKUDA M,et al.Izumoring:A novel and complete strategy for bioproduction of rare sugars[J].Journal of Bioscience and Bioengineering,2004,97(2):89-94.
[2] CHATTOPADHYAY S,RAYCHAUDHURI U,CHAKRABORTY R.Artificial sweeteners-A review[J].Journal of Food Science and Technology,2014,51(4):611-621.
[3] KIMURA T,KANASAKI A,HAYASHI N,et al.D-allulose enhances postprandial fat oxidation in healthy humans[J].Nutrition,2017,43-44:16-20.
[4] IWASAKI Y,SENDO M,DEZAKI K,et al.GLP-1 release and vagal afferent activation mediate the beneficial metabolic and chronotherapeutic effects of D-allulose[J].Nature Communications,2018,9:113.
[5] SUNA S,YAMAGUCHI F,KIMURA S,et al.Preventive effect of D-psicose,one of rare ketohexoses,on di-(2-ethylhexyl) phthalate (DEHP)-induced testicular injury in rat[J].Toxicology Letters,2007,173(2):107-117.
[6] TAKATA M K,YAMAGUCHI F,NAKANOSE K,et al.Neuroprotective effect of D-psicose on 6-hydroxydopamine-induced apoptosis in rat pheochromocytoma (PC12) cells[J].Journal of Bioscience and Bioengineering,2005,100(5):511-516.
[7] ANDREANA P R,MCLELLAN J S,CHEN Y,et al.Synthesis of 2,6-dideoxysugars via ring-closing olefinic metathesis[J].Organic Letters,2002,4(22):3 875-3 878.
[8] NORTHRUP A B,MACMILLAN D W C.Two-step synthesis of carbohydrates by selective aldol reactions[J].Cheminform,2004,5691:1 752-1 754.
[9] DONER L W.Isomerization of D-fructose by base:Liquid-chromatographic evaluation and the isolation of D-psicose[J].Carbohydrate Research,1979,70(2):209-216.
[10] SOENGAS R,IZUMORI K,SIMONE M I,et al.Kiliani reactions on ketoses:Branched carbohydrate building blocks from D-tagatose and D-psicose[J].Tetrahedron Letters,2005,46(34):5 755-5 759.
[11] 方志杰, 李松,程杰,等.一种由糖酸内酯出发合成稀有己酮糖和庚酮糖的方法:中国,200910031931.7[P].2009-07-03.
FANG Z J,LI S,CHENG J,et al.A method for the synthesis of rare hexoses and heptanoses from glycoacid lactones:China,200910031931.7[P].2009-07-03.
[12] IZUMORI K,KHAN A R,OKAYA H,et al.A new enzyme,D-ketohexose 3-Epimerase,from Pseudomonas sp.ST-24[J].Bioscience Biotechnology and Biochemistry,1993,57(6):1 037-1 039.
[13] KIM H J,HYUN E K,KIM Y S,et al.Characterization of an Agrobacterium tumefaciens D-psicose 3-Epimerase that converts D-fructose to D-psicose[J].Applied and Environmental Microbiology,2006,72(2):981-985.
[14] ZHU Y M,MEN Y,BAI W,et al.Overexpression of D-psicose 3-epimerase from Ruminococcus sp.in Escherichia coli and its potential application in D-psicose production[J].Biotechnology Letters,2012,34(10):1 901-1 906.
[15] JIA M,MU W M,CHU F F,et al.A D-psicose 3-epimerase with neutral pH optimum from Clostridium bolteae for D-psicose production:Cloning,expression,purification,and characterization[J].Applied Microbiology and Biotechnology,2014,98(2):717-725.
[16] YOSHIHARA A,KOZAKAI T,SHINTANI T,et al.Purification and characterization of D-allulose 3-epimerase derived from Arthrobacter globiformis M30,a GRAS microorganism[J].Journal of Bioscience and Bioengineering,2017,123(2):170-176.
[17] ZHANG W L,LI H,ZHANG T,et al.Characterization of a D-psicose 3-epimerase from Dorea sp.CAG317 with an acidic pH optimum and a high specific activity[J].Journal of Molecular Catalysis B:Enzymatic,2015,120:68-74.
[18] ZHANG W L,ZHANG T,JIANG B,et al.Biochemical characterization of a D-psicose 3-epimerase from Treponema primitia ZAS-1 and its application on enzymatic production of D-psicose[J].Journal of the Science of Food and Agriculture,2016,96(1):49-56.
[19] MU W M,CHU F F,XING Q C,et al.Cloning,expression,and characterization of a D-psicose 3-Epimerase from Clostridium cellulolyticum H10[J].Journal of Agricultural and Food Chemistry,2011,59(14):7 785-7 792.
[20] ZHANG W L,FANG D,ZHANG T,et al.Characterization of a metal-dependent D-psicose 3-epimerase from a novel strain,Desmospora sp.8437[J].Journal of Agricultural and Food Chemistry,2013,61(47):11 468-11 476.
[21] MU W M,ZHANG W L,FANG D,et al.Characterization of a D-psicose-producing enzyme,D-psicose 3-epimerase,from Clostridium sp.[J].Biotechnology Letters,2013,35(9):1 481-1 486.
[22] ZHANG W L,FANG D,XING Q C,et al.Characterization of a novel metal-dependent D-psicose 3-epimerase from Clostridium scindens 35704[J].PLoS ONE,2013,8(4):e62 987.
[23] PARK C,KIM T,HONG S,et al.D-allulose production from D-fructose by permeabilized recombinant cells of Corynebacterium glutamicum cells expressing D-allulose 3-epimerase Flavonifractor plautii[J].PLoS One,2016,11(7):e160 044.
[24] KIM K,KIM H,OH D,et al.Crystal structure of D-psicose 3-epimerase from Agrobacterium tumefaciens and its complex with true substrate D-fructose:A pivotal role of metal in catalysis,an active site for the non-phosphorylated substrate,and its conformational changes[J].Journal of Molecular Biology,2006,361(5):920-931.
[25] QI Z L,ZHU Z L,WANG J W,et al.Biochemical analysis and the preliminary crystallographic characterization of D-tagatose 3-epimerase from Rhodobacter sphaeroides[J].Microbial Cell Factories,2017,16(1):193.
[26] KIM H,LIM B,YEOM S,et al.Roles of Ile66 and Ala107 of D-psicose 3-epimerase from Agrobacterium tumefaciens in binding O6 of its substrate,D-fructose[J].Biotechnology Letters,2010,32(1):113-118.
[27] 张黎丽,朱玥明,门燕,等.来源于瘤胃菌Ruminococcus sp.的D-阿洛酮糖3-差向异构酶的底物结合位点分析[J].微生物学通报,2014,41(5):811-817.
ZHANG L L,ZHU Y M,MEN Y,et al.Substrate-binding site of D-psicose 3-epimerase from Ruminococcus sp.[J].Microbiology China,2014,41(5):811-817.
[28] ZHU Z L,GAO D K,LI C,et al.Redesign of a novel D-allulose 3-epimerase from Staphylococcus aureus for thermostability and efficient biocatalytic production of D-allulose[J].Microbial Cell Factories,2019,18(1):59.
[29] ZHU J X,LI Y,WANG J Z,et al.Adaptive steered molecular dynamics combined with protein structure networks revealing the mechanism of Y68I/G109P mutations that enhance the catalytic activity of D-psicose 3-epimerase from Clostridium bolteae[J].Frontiers in Chemistry,2018,6:437.
[30] TOLMAN R C.The temperature coefficient of photochemical reaction rate[J].Journal of the American Chemical Society,1923,45(10):2 285-2 296.
[31] CHOI J,JU Y,YEOM S,et al.Improvement in the thermostability of D-psicose 3-epimerase from Agrobacterium tumefaciens by random and site-directed mutagenesis[J].Applied and Environmental Microbiology,2011,77(20):7 316-7 320.
[32] ZHANG W L,JIA M,YU S H,et al.Improving the thermostability and catalytic efficiency of the d-psicose 3-epimerase from Clostridium bolteae ATCC BAA-613 using site-directed mutagenesis[J].Journal of Agricultural and Food Chemistry,2016,64(17):3 386-3 393.
[33] ZHANG W L,ZHANG Y M,HUANG J W,et al.Thermostability improvement of the D-allulose 3-epimerase from Dorea sp.CAG317 by site-directed mutagenesis at the interface regions[J].Journal of Agricultural and Food Chemistry,2018,66(22):5 593-5 601.
[34] MAO S H,CHENG X T,ZHU Z L,et al.Engineering a thermostable version of D-allulose 3-epimerase from Rhodopirellula baltica via site-directed mutagenesis based on B-factors analysis[J].Enzyme and Microbial Technology,2020,132:109 441.
[35] 谢雪原, 周华,高震,等.多酶偶联反应体系的研究和应用[J].化工进展,2004,23(8):864-868.
XIE X Y,ZHOU H,GAO Z,et al.Study and application of coupled multienzyme reaction system[J].Chemical Industry and Engineering Progress,2004,23(8):864-868.
[36] MEN Y,ZHU Y M,ZENG Y,et al.Co-expression of D-glucose isomerase and D-psicose 3-epimerase:Development of an efficient one-step production of D-psicose[J].Enzyme and Microbial Technology,2014,64-65:1-5.
[37] 高雅慧, 金则成,钱超,等.基于RhaD醛缩酶的“一锅四酶法”合成D-山梨糖和D-阿洛酮糖[J].食品工业科技,2013,34(21):210-214.
GAO Y H,JIN Z C,QIAN C,et al.One pot four enzyme synthesis of D-sorbose and D-psicose based on aldolase RhaD[J].Science and Technology of Food Industry,2013,34(21):210-214.
[38] 韩文佳, 朱玥明,柏玮,等.双酶偶联转化果糖制备含有稀少糖的混合糖液[J].生物工程学报,2014,30(1):90-97.
HAN W J,ZHU Y M,BO W et al.Production of sugar syrup containing rare sugar using dual-enzyme coupled reaction system[J].Chinese Journal of Biotechnology,2014,30(1):90-97.
[39] 贾敏, 沐万孟,张涛,等.D-阿洛酮糖3-差向异构酶基因在枯草芽孢杆菌中的表达[J].食品与生物技术学报,2014,33(11):1 129-1 135.
JIA M,MU W M,ZHANG T,et al.Expression of D-psicose 3-epimerase in Bacillus subtilis[J].Journal of Food Science and Biotechnology,2014,33(11):1 129-1 135.
[40] HE W W,JIANG B,MU W M,et al.Production of D-allulose with D-psicose 3-epimerase expressed and displayed on the surface of Bacillus subtilis spores[J].Journal of Agricultural and Food Chemistry,2016,64(38):7 201-7207
[41] CHEN J Q,ZHAO L Q,FU G,et al.A novel strategy for protein production using non-classical secretion pathway in Bacillus subtilis[J].Microbial Cell Factories,2016,15(1):1-16.
[42] YANG J G,TIAN C Y,ZHANG T,et al.Development of food-grade expression system for D-allulose 3-epimerase preparation with tandem isoenzyme genes in Corynebacterium glutamicum and its application in conversion of cane molasses to D-allulose[J].Biotechnology and Bioengineering,2019,116(4):745-756.
[43] LIM B,KIM H,OH D.A stable immobilized D-psicose 3-epimerase for the production of D-psicose in the presence of borate[J].Process Biochemistry,2009,44(8):822-828.
[44] YOSHIHARA A,KOZAKAI T,SHINTANI T,et al.Purification and characterization of D-allulose 3-epimerase derived from Arthrobacter globiformis M30,a GRAS microorganism[J].Journal of Bioscience and Bioengineering,2017,123(2):170-176.
[45] TSENG C,LIAO C,SUN Y,et al.Immobilization of Clostridium cellulolyticum D-psicose 3-epimerase on artificial oil bodies[J].Journal of Agricultural and Food Chemistry,2014,62(28):6 771-6 776.
[46] LI C,LIN J Q,GUO Q Q,et al.D-psicose 3-epimerase secretory overexpression,immobilization,and D-psicose biotransformation,separation and crystallization[J].Journal of Chemical Technology & Biotechnology,2018,93(2):350-357.
[47] 孙帆, 宿玲恰,张康,等.D-阿洛酮糖3-差向异构酶在枯草芽孢杆菌中的高效表达及固定化细胞研究[J].中国生物工程杂志,2018,38(7):83-88.
SUN F,SU L Q,ZHANG K,et al.D-psicose 3-epimerase gene overexpression in Bacillus subtilis and immobilization of cells[J].China Biotechnology,2018,38(7):83-88.
[48] NARAYAN PATEL S,SINGH V,SHARMA M,et al.Development of a thermo-stable and recyclable magnetic nanobiocatalyst for bioprocessing of fruit processing residues and D-allulose synthesis[J].Bioresource Technology,2018,247:633-639.
[49] ZHANG X J,FAN H H,LIU N,et al.A novel self-sufficient biocatalyst based on transaminase and pyridoxal 5′-phosphate covalent co-immobilization and its application in continuous biosynthesis of sitagliptin[J].Enzyme and Microbial Technology,2019,130:109 362.
[50] 金利群, 郭东京,廖承军.固定化葡萄糖异构酶的研究进展[J].发酵科技通讯,2015,44(3):47-51.
JIN L Q,GUO D J,LIAO C J.Progress in immobilized glucose isomerase[J].Bulletin of Fermentation Science and Technology,2015,44(3):47-51.