[1] 包启安. 酱油科学与酿造技术[M].北京:中国轻工业出版社, 2011:44-45.
[2] FENG Y Z, SU G W, ZHAO H F, et al.Characterisation of aroma profiles of commercial soy sauce by odour activity value and omission test[J].Food Chemistry, 2015, 167:220-228.
[3] DEVANTHI P V P, LINFORTH R, ONYEAKA H, et al.Effects of co-inoculation and sequential inoculation of Tetragenococcus halophilus and Zygosaccharomyces rouxii on soy sauce fermentation[J].Food Chemistry, 2018, 240:1-8.
[4] DEVANTHI P V P, GKATZIONIS K.Soy sauce fermentation:Microorganisms, aroma formation, and process modification[J].Food Research International, 2019, 120:364-374.
[5] ZHAO G Z, DING L L, PAN Z H, et al.Proteinase and glycoside hydrolase production is enhanced in solid-state fermentation by manipulating the carbon and nitrogen fluxes in Aspergillus oryzae[J].Food Chemistry, 2019, 271:606-613.
[6] ZHAO G Z, LIU C, LI S, et al.Exploring the flavor formation mechanism under osmotic conditions during soy sauce fermentation in Aspergillus oryzae by proteomic analysis[J].Food & Function, 2020, 11(1):640-648.
[7] 谭戈, 徐晴元, 郭丽琼, 等.酿造酱油微生物多样性及风味物质研究进展[J].中国调味品, 2021, 46(6):160-163;179.
TAN G, XU Q Y, GUO L Q, et al.Research progress on the microbial diversity and flavor substances in brewed soy sauce[J].China Condiment, 2021, 46(6):160-163;179.
[8] SULAIMAN J, GAN H M, YIN W F, et al.Microbial succession and the functional potential during the fermentation of Chinese soy sauce brine[J].Frontiers in Microbiology, 2014, 5:556.
[9] HARADA R, YUZUKI M, ITO K, et al.Microbe participation in aroma production during soy sauce fermentation[J].Journal of Bioscience and Bioengineering, 2018, 125(6):688-694.
[10] HARADA R, YUZUKI M, ITO K, et al.Influence of yeast and lactic acid bacterium on the constituent profile of soy sauce during fermentation[J].Journal of Bioscience and Bioengineering, 2017, 123(2):203-208.
[11] QI Q, HUANG J, ZHOU R Q, et al.Characterizing microbial community and metabolites of Cantonese soy sauce[J].Food Bioscience, 2021, 40:100872.
[12] YANG Y, DENG Y E, JIN Y L, et al.Dynamics of microbial community during the extremely long-term fermentation process of a traditional soy sauce[J].Journal of the Science of Food and Agriculture, 2017, 97(10):3220-3227.
[13] 冯云子. 高盐稀态酱油关键香气物质的变化规律及形成机理的研究[D].广州:华南理工大学,2015.
FENG Y Z.The evolution and formation mechanism of key aroma compounds during the process of high-salt liquid fermentation soy Sauce[D].Guangzhou:South China University of Technology, 2015.
[14] DIEZ-SIMON C, EICHELSHEIM C, MUMM R, et al.Chemical and sensory characteristics of soy sauce:A review[J].Journal of Agricultural and Food Chemistry, 2020, 68(42):11612-11630.
[15] 陈杰, 赵莹, 韩舜羽, 等.17种市售广式酱油中风味物质的检测分析[J].中国酿造, 2021, 40(12):165-170.
CHEN J, ZHAO Y, HAN S Y, et al.Detection and analysis of flavor substances in 17 kinds of commercially available Cantonese soy sauce[J].China Brewing, 2021, 40(12):165-170.
[16] 刘非. 基于GC-MS的酱油风味物质分析[D].天津:天津科技大学, 2017.
LIN F.Analysis of aroma compounds of soy sauce based on GC-MS[D].Tianjin:Tianjin University of Science and Technology, 2017.
[17] ZHAO G Z, DING L L, HADIATULLAH H, et al.Characterization of the typical fragrant compounds in traditional Chinese-type soy sauce[J].Food Chemistry, 2020, 312:126054.
[18] 朱新贵, 李学伟, 曾小波.典型广式酱油与日式酱油的风味物质差异研究[J].中国酿造, 2016, 35(7):30-35.
ZHU X G, LI X W, ZENG X B.Difference of flavor compounds in Cantonese-style soy sauce and Japanese-style soy sauce[J].China Brewing, 2016, 35(7):30-35.
[19] 白佳伟, 陈亮, 周尚庭, 等.特级高盐稀态酿造酱油中关键香气物质的分析[J].中国酿造, 2019, 38(11):179-185.
BAI J W, CHEN L, ZHOU S T, et al.Analysis of key odor compounds in super grade soy sauce with high-salt and diluted-state fermentation[J].China Brewing, 2019, 38(11):179-185.
[20] 阮志强, 董玺梅, 蒋雪薇, 等.高盐稀态酱油发酵优势真菌与风味物质相关性分析[J].食品科学, 2022, 43(10):172-179.
RUAN Z Q, DONG X M, JIANG X W, et al.Correlation between dominant fungi and variation of flavor compounds during high-salt liquid-state soy sauce fermentation[J].Food Science, 2022, 43(10):172-179.
[21] ZHAO G Z, DING L L, YAO Y P, et al.Extracellular proteome analysis and flavor formation during soy sauce fermentation[J].Frontiers in Microbiology, 2018, 9:1872.
[22] SUN S Y, JIANG W G, ZHAO Y P.Profile of volatile compounds in 12 Chinese soy sauces produced by a high-salt-diluted state fermentation[J].Journal of the Institute of Brewing, 2010, 116(3):316-328.
[23] DZIALO M C, PARK R, STEENSELS J, et al.Physiology, ecology and industrial applications of aroma formation in yeast[J].FEMS Microbiology Reviews, 2017, 41(S1):S95-S128.
[24] ALBA-LOIS L,SEGAL-KISCHINEVZKY C.Yeast fermentation and the making of beer and wine[J].Nature Education, 2010, 3(9):17.
[25] AERVEAANSKAYĞ I, MIHAL M, MARKO J.Potential application of perfusion and pertraction for in situ product removal in biocatalytic 2-phenylethanol production[J].Separation and Purification Technology, 2017, 183:11-20.
[26] FENG Y Z, CAI Y, SU G W, et al.Evaluation of aroma differences between high-salt liquid-state fermentation and low-salt solid-state fermentation soy sauces from China[J].Food Chemistry, 2014, 145:126-134.
[27] GAO X L, CUI C, REN J Y, et al.Changes in the chemical composition of traditional Chinese-type soy sauce at different stages of manufacture and its relation to taste[J].International Journal of Food Science & Technology, 2011, 46(2):243-249.
[28] 冯云子, 周婷, 吴伟宇, 等.酱油风味与功能性成分研究进展[J].食品科学技术学报, 2021, 39(4):14-28.
FENG Y Z, ZHOU T, WU W Y, et al.Research progress on flavor and functional components of soy sauce[J].Journal of Food Science and Technology, 2021, 39(4):14-28.
[29] 赵谋明, 许瑜, 苏国万, 等.不同淀粉质原料对高盐稀态酱油香气品质的影响[J].现代食品科技, 2018, 34(6):130-142.
ZHAO M M, XU Y, SU G W, et al.Effect of different starchy materials on aroma quality of high-salt dilute soy sauce[J].Modern Food Science and Technology, 2018, 34(6):130-142.
[30] RAMASWAMY H S, RICHARDS J F.Flavor of poultry meat:A review[J].Canadian Institute of Food Science and Technology Journal, 1982, 15(1):7-18.
[31] ZHANG Y, SONG H L, LI P, et al.Determination of potential off-flavour in yeast extract[J].LWT - Food Science and Technology, 2017, 82:184-191.
[32] 吴拥军. 酱(豉)香风味的研究进展[J].山地农业生物学报, 2018, 37(6):1-9.
WU Y J.Research porgress of Maotai flanor[J].Journal of Mountain Agriculture and Biology, 2018, 37(6):1-9.
[33] SUGAWARA E, HASHIMOTO S, SAKURAI Y, et al.Formation by yeast of the HEMF (4-hydroxy-2(or 5)-ethyl-5(or 2)-methyl-3(2H)-furanone) aroma component in miso with aging[J].Bioscience, Biotechnology, and Biochemistry, 1994, 58(6):1134-1135.
[34] OHATA M, KOHAMA K, MORIMITSU Y, et al.The formation mechanism by yeast of 4-hydroxy-2(or 5)-ethyl-5(or 2)-methyl-3(2H)-furanone in miso[J].Bioscience, Biotechnology, and Biochemistry, 2007, 71(2):407-413.
[35] LIU S Q, HOLLAND R, CROW V L.Esters and their biosynthesis in fermented dairy products:A review[J].International Dairy Journal, 2004, 14(11):923-945.
[36] ZHANG L J, ZHANG L, XU Y.Effects of Tetragenococcus halophilus and Candida versatilis on the production of aroma-active and umami-taste compounds during soy sauce fermentation[J].Journal of the Science of Food and Agriculture, 2020, 100(6):2782-2790.
[37] KANEKO S, KUMAZAWA K, NISHIMURA O.Studies on the key aroma compounds in raw (unheated) and heated Japanese soy sauce[J].Journal of Agricultural and Food Chemistry, 2013, 61(14):3396-3402.
[38] 高献礼, 赵谋明, 崔春, 等.高盐稀态酱油挥发性风味物质的分离与鉴定[J].华南理工大学学报(自然科学版), 2009, 37(10):117-123.
GAO X L, ZHAO M M, CUI C, et al.Separation and identification of volatile flavor compounds in high-salt liquid state-fermented soy sauce[J].Journal of South China University of Technology (Natural Science Edition), 2009, 37(10):117-123.
[39] 崔云前, 曹小红, 王春玲, 等.发酵行业4-乙烯基愈创木酚和4-乙基愈创木酚研究进展[J].中国酿造, 2009, 28(4):14-17.
CUI Y Q, CAO X H, WANG C L, et al.Research progress of 4-vinylguaiacol and 4-ethylguaiacol in fermentation industry[J].China Brewing, 2009, 28(4):14-17.
[40] ITO T, KONNO M, SHIMURA Y, et al.Formation of guaiacol by spoilage bacteria from vanillic acid, a product of rice koji cultivation, in Japanese sake brewing[J].Journal of Agricultural and Food Chemistry, 2016, 64(22):4599-4605.
[41] SUEZAWA Y, SUZUKI M.Bioconversion of ferulic acid to 4-vinylguaiacol and 4-ethylguaiacol and of 4-vinylguaiacol to 4-ethylguaiacol by halotolerant yeasts belonging to the genus Candida[J].Bioscience, Biotechnology, and Biochemistry, 2007, 71(4):1058-1062.
[42] MUIR D D.The Maillard Reaction? Chemistry, biochemistry and implications[J].International Journal of Dairy Technology, 2007, 60(1):59.
[43] 张怀志. 重要风味化合物2-乙基-3,5/3,6-二甲基吡嗪的微生物合成途径解析[D].无锡:江南大学, 2020.
ZHANG H Z.The bio-synthetic pathway of 2-ethyl-3,5/3,6-dimethylpyrazine, the valuable flavor compounds[D].Wuxi:Jiangnan University, 2020.
[44] CERNY C.The aroma side of the Maillard reaction[J].Annals of the New York Academy of Sciences, 2008, 1126(1):66-71.
[45] WEI Q Z, WANG H B, CHEN Z X, et al.Profiling of dynamic changes in the microbial community during the soy sauce fermentation process[J].Applied Microbiology and Biotechnology, 2013, 97(20):9111-9119.
[46] SINGRACHA P, NIAMSIRI N, VISESSANGUAN W, et al.Application of lactic acid bacteria and yeasts as starter cultures for reduced-salt soy sauce (moromi) fermentation[J].LWT, 2017, 78:181-188.
[47] 邹谋勇, 朱新贵, 曾小波, 等.一株接合酵母FW30-2及其应用:中国, CN110408555B[P].2021-04-23.
ZOU MY, ZHU X G, ZENG X B, et al.A strain of Zygosaccharomyces sp FW30-2 and its application:China, CN110408555B[P].2021-04-23.
[48] DAKAL T C, SOLIERI L, GIUDICI P.Adaptive response and tolerance to sugar and salt stress in the food yeast Zygosaccharomyces rouxii[J].International Journal of Food Microbiology, 2014, 185:140-157.
[49] CAO Z H, GREEN-JOHNSON J M, BUCKLEY N D, et al.Bioactivity of soy-based fermented foods:A review[J].Biotechnology Advances, 2019, 37(1):223-238.
[50] JANSEN M, VEURINK J H, EUVERINK G J W, et al.Growth of the salt-tolerant yeast Zygosaccharomyces rouxii in microtiter plates:Effects of NaCl, pH and temperature on growth and fusel alcohol production from branched-chain amino acids[J].FEMS Yeast Research, 2003, 3(3):313-318.
[51] YAO S J, ZHOU R Q, JIN Y, et al.Effect of co-culture with Tetragenococcus halophilus on the physiological characterization and transcription profiling of Zygosaccharomyces rouxii[J].Food Research International, 2019, 121:348-358.
[52] 方冠宇, 姜佳丽, 蒋予箭.多菌混合发酵对酱油的风味物质形成及感官指标的影响[J].中国食品学报, 2019, 19(9):154-163.
FANG G Y, JIANG J L, JIANG Y J.The influence of multi-strain fermentation on the flavor substances and sensory evaluation in soy sauce[J].Journal of Chinese Institute of Food Science and Technology, 2019, 19(9):154-163.
[53] HAUCK T, BRÜHLMANN F, SCHWAB W.Formation of 4-hydroxy-2, 5-dimethyl-3[2H]-furanone by Zygosaccharomyces rouxii:Identification of an intermediate[J].Applied and Environmental Microbiology, 2003, 69(7):3911-3918.
[54] 谭戈, 朱新贵, 李学伟, 等.强化鲁氏接合酵母对酱油品质的影响[J].食品工业科技, 2022, 43(13):86-93.
TAN G, ZHU X G, LI X W, et al.Effect of fortified Zygosaccharomyces rouxii on the quality of soy sauce[J].Science and Technology of Food Industry, 2022, 43(13):86-93.
[55] 许焰, 吴群, 徐岩.酱香型白酒酿造主要功能菌株对拜耳接合酵母的作用[J].微生物学通报, 2018, 45(1):42-53.
XU Y, WU Q, XU Y.Effects of main functional strains on Zygosaccharomyces bailii in Chinese Maotai-flavor liquor fermentation[J].Microbiology China, 2018, 45(1):42-53.
[56] 周钰涵, 崔丹瑶, 闫昕, 等.酿酒酵母与球拟酵母混合发酵对白酒风味物质的影响[J].食品研究与开发, 2021, 42(17):169-175.
ZHOU Y H, CUI D Y, YAN X, et al.Effect of mixed fermentation of Saccharomyces cerevisiae and Torulopsis globosa on flavor substances of Baijiu[J].Food Research and Development, 2021, 42(17):169-175.
[57] WANAKHACHORNKRAI P.Comparison of determination method for volatile compounds in Thai soy sauce[J].Food Chemistry, 2003, 83(4):619-629.
[58] 郭建, 伍学明, 樊君, 等.鲁氏酵母和球拟酵母不同接种方式对高盐稀态酿造酱油品质的影响[J].中国调味品, 2019, 44(2):100-104.
GUO J, WU X M, FAN J, et al.Effect of different inoculation modes with Saccharomyces rouxii and Torulopsis bombicola on the quality of soy sauce by high-salt and liquid-state fermentation[J].China Condiment, 2019, 44(2):100-104.
[59] 张丽. 高盐稀态酱油发酵过程中添加酵母的研究[D].贵阳:贵州大学, 2019.
ZHANG L.Study on adding yeast in the fermentation process of high-salt dilute soy sauce[D].Guiyang:Guizhou University, 2019.
[60] HIGUCHI T, AOKI T, UCHIDA K.Esterase activity in soy sauce moromi as a factor hydrolyzing flavor esters[J].Journal of Fermentation and Bioengineering, 1991, 71(3):163-167.
[61] LIU B, LI Y, CAO Z N, et al.Effect of Tetragenococcus halophilus, Zygosaccharomyces rouxii, and Torulopsis versatilis addition sequence on soy sauce fermentation[J].Innovative Food Science & Emerging Technologies, 2021, 69:102662.
[62] YAN Y Z, QIAN Y L, JI F D, et al.Microbial composition during Chinese soy sauce koji-making based on culture dependent and independent methods[J].Food Microbiology, 2013, 34(1):189-195.
[63] QI W, GUO H L, WANG C L, et al.Comparative study on fermentation performance in the genome shuffledCandida versatilisand wild-type salt tolerant yeast strain[J].Journal of the Science of Food and Agriculture, 2017, 97(1):284-290.
[64] 张玲. 嗜盐四联球菌、鲁氏接合酵母及易变假丝酵母的添加对日式酱油品质的影响[D].无锡:江南大学, 2019.
ZHANG L.Effect of Tetragenococcus halophilus, Zygosaccharomyces rouxii and Candida versatilis on Japanese soy sauce quality[D].Wuxi:Jiangnan University, 2019.
[65] 邹谋勇, 朱新贵, 李学伟, 等.假丝酵母菌FW922-1及其应用:中国, CN109370927B[P].2021-06-01.
ZOU M Y, ZHU X G, LI X W, et al.Candida FW922-1 and its application:China, CN109370927B[P].2021-06-01.
[66] 韩冉, 张玲玲, 黄文烨, 等.假丝酵母的添加对高盐稀态酱油有机酸含量影响[J].中国调味品, 2021, 46(5):5-9.
HAN R, ZHANG L L, HUANG W Y, et al.Effect of Candida on organic acid content in high salt dilute soy sauce[J].China Condiment, 2021, 46(5):5-9.
[67] WANG J W, ZHAO M M, XIE N Y, et al.Community structure of yeast in fermented soy sauce and screening of functional yeast with potential to enhance the soy sauce flavor[J].International Journal of Food Microbiology, 2022, 370:109652.
[68] 冯杰. 埃切假丝酵母产香机理及其对酱油风味的影响[D].无锡:江南大学, 2012.
FENG J.Mechanism of aroma production for the improvement of soy sauce flavor by Candida etchellsii[D].Wuxi: Jiangnan University, 2012.
[69] SONG Y R, JEONG D Y, BAIK S H.Monitoring of yeast communities and volatile flavor changes during traditional Korean soy sauce fermentation[J].Journal of Food Science, 2015, 80(9):M2005-M2014.
[70] KHOT M, GHOSH D.Lipids of Rhodotorula mucilaginosa IIPL32 with biodiesel potential:Oil yield, fatty acid profile, fuel properties[J].Journal of Basic Microbiology, 2017, 57(4):345-352.
[71] WAH T T, WALAISRI S, ASSAVANIG A, et al.Co-culturing of Pichia guilliermondii enhanced volatile flavor compound formation by Zygosaccharomyces rouxii in the model system of Thai soy sauce fermentation[J].International Journal of Food Microbiology, 2013, 160(3):282-289.
[72] CAO X H, SONG Q, WANG C L, et al.Genome shuffling of Hansenula anomala to improve flavour formation of soy sauce[J].World Journal of Microbiology and Biotechnology, 2012, 28(5):1857-1862.
[73] TANAKA Y, WATANABE J, MOGI Y.Monitoring of the microbial communities involved in the soy sauce manufacturing process by PCR-denaturing gradient gel electrophoresis[J].Food Microbiology, 2012, 31(1):100-106.
[74] VALACHOVÁ K, HORVÁTHOVÁ V.Starch degradation by glucoamylase Glm from Saccharomycopsis fibuligera IFO 0111 in the presence and absence of a commercial pullulanase[J].Chemistry & Biodiversity, 2007, 4(5):874-880.